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I plan on draping the cage with a towel or piece of fleece to make it a little more closed and safe-feeling for them, is this a good idea?
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If it's cool enough in the vehicle you can do this to make them feel more secure but if it's hot or you don't plan on running the air conditioning, I'm not sure if I'd do it.
Is there anyway you could provide 2 traveling cages as one really isn't going to provide much room for 4. I know it would take up more room but it might help to prevent fights or arguments plus give them a little more room to move around in. Maybe you could use your one pet store cage and then make a 1x2 C&C as well. Another reason to provide an extra traveling cage is to have room for each a hidey house. That way they'd have a place to hide and would be more at ease and cut down on stress.
Also provide plenty of fresh wet veggies along the way to ensure no one gets dehydrated and give plenty of hay in the cage(s). Hanging a water bottle in a moving car is hard because many bottles tend to leak due to the movement of the vehicle. Take a water bottle along though and stop frequently to give them drinks from it.